bettle

Übersetzung im Kontext von „Ich bettle“ in Deutsch-Englisch von Reverso Context: Möchtest du, dass ich bettle?. Präsens. ich bettle du bettelst er/sie/es bettelt wir betteln ihr bettelt sie/Sie betteln. Präteritum. ich bettelte bettle betteln wir bettelt betteln Sie. Infinitiv. Präsens. Seit Oktober sind Frau Dr. Anke Ribbeck und seit Juli Herr Herr Dr. Ingo Görzig neben Herrn Dr. Oliver Bettle Mitinhaber der Praxis, zu der 13 weitere.

The bodywork attached with eighteen bolts to its nearly flat platform chassis which featured a central structural tunnel. Front and rear suspension featured torsion bars along with front stabilizer bar—providing independent suspension at all wheels.

Certain initial features were subsequently revised, including mechanical drum brakes, split-window rear windows, mechanical direction-indicators and the non-synchronized gearbox.

Other features, including its distinctive overall shape, endured. In fact, the Beetle was prized for its seemingly unchanged appearance and "marketed to American consumers as the anti-GM and Ford: Its engine, transmission, and cylinder heads were constructed of light alloy.

Later models of the carburetor featured an automatic choke. Engine intake air passed through a metallic filter, while heavier particles were captured by an oil bath.

After , steering featured a hydraulic damper that absorbed steering irregularities. Throughout its production, VW marketed the Beetle with a four-speed manual transmission.

From and almost exclusively in Europe , [54] VW offered an optional version of the Saxomat semi-automatic transmission: Subsequently, beginning in in Europe and in the United States , VW offered an optional semi-automatic transmission marketed as Automatic Stick Shift and also called AutoStick , [ citation needed ] which was a 3-speed manual coupled to an electro-pneumatic clutch and torque converter.

It was in that Wilhelm Karmann first bought a VW Beetle sedan and converted it into a four-seated convertible. The convertible was more than a Beetle with a folding top.

To compensate for the strength lost in removing the roof, the sills were reinforced with welded U-channel rails, a transverse beam was fitted below the front edge of the rear seat cushion, and the side cowl-panels below the instrument panel were double-wall.

In addition, the lower corners of the door apertures had welded-in curved gussets, and the doors had secondary alignment wedges at the B-pillar.

The top was cabriolet-style with a full inner headliner hiding the folding mechanism and crossbars. The rear window was tempered safety glass, and after , heated.

Due to the thickness of the top, it remained quite tall when folded. To enable the driver to see over the lowered top, the inside rearview was mounted on an offset pivot.

The convertible was generally more lavishly equipped than the sedan with dual rear ashtrays, twin map pockets, a visor vanity mirror on the passenger side, rear stone shields, and through , wheel trim rings.

Many of these items did not become available on other Beetles until the advent of the optional "L" Luxus Package of After a number of stylistic and technical alterations made to the Karmann cabriolet, [57] corresponding to the many changes VW made to the Beetle throughout its history , the last of , cabriolets came off the production line on 10 January During this period, a myriad of changes were made throughout the vehicle beginning with the availability of hydraulic brakes and a folding fabric sunroof in The rear window of the VW Beetle evolved from a divided or "split" oval, to a singular oval.

The change occurred between October and March The brake light function was transferred to new heart-shaped lamps located in the top of the taillight housings.

In , the separate brake lights were discontinued and were combined into a new larger taillight housing.

The traditional VW semaphore turn signals were replaced by conventional flashing directional indicator lamps for North America.

For , the Beetle received what would become one of its more distinctive features, a set of twin chrome tailpipes. Models for North America gained taller bumper guards and tubular overrider bars.

For , the Beetle received a revised instrument panel, and a larger rectangular rear window replaced the previous oval design. For , significant technical advances occurred in the form of a new engine and transmission.

The single-barrel Solex carburetor received an electric automatic choke and the transmission was now synchronized on all forward gears.

The traditional semaphore turn signals were replaced by conventional flashing directional indicators worldwide.

For , the Beetle received a mechanical fuel level gauge in place of the former fuel tap. The Standard model continued without a gas gauge until the end of the model year.

At the rear, larger tail lights were introduced incorporating a separate amber turn signal section to meet new European standards these turn signals remained red in the US market until The former hand-pump style windscreen washer was replaced by a new design using compressed air.

The largest change to date for the Beetle was in The result was a more open, airy, modern look. Models so equipped carried a "" badge on the engine lid.

The cc engine was standard for North America. For , a yet-again larger- displacement engine was made available: North America received the engine as standard equipment, but did not receive front disc brakes.

These models were identified by a "Volkswagen" badge on the engine lid. The rear suspension was significantly revised and included a widened track, softer torsion bars and the addition of a unique Z-configuration equalizing torsion spring.

The clutch disc also increased in size and changes were made to the flywheel. In accord with the newly enacted US Federal Motor Vehicle Safety Standard , North American models received a dual-circuit brake system, the clear glass headlamp covers were deleted; the headlamps were brought forward to the leading edge of the front fenders, and the sealed-beam units were exposed and surrounded by chrome bezels.

In the rest of the world markets the model retained the older headlights. Another oddity of the North American market Beetle is the rear bumper overriders towel rails — the overriders have a different shape than the older models besides the one-year only engine decklid.

The most noticeable of which were the new larger, higher mounted C-section bumpers. At the rear, new larger taillamps were adopted and were able to accommodate reversing lamps, which were previously separate bumper-mounted units.

Other improvements were a new outside gas filler with spring-loaded flap, eliminating the need to open the trunk to refuel. The fuel gauge was integrated with the speedometer and was now electrically actuated rather than cable-operated.

A number of safety improvements were made in order to comply with new American safety regulations: US models received a padded instrument panel that was optional in other markets.

A new 3-speed semi-automatic gear box with torque converter and vacuum-operated clutch became available mid-production year.

The semi-automatic models received a vastly improved semi-trailing-arm rear suspension also known as "independent rear suspension" although the earlier swing axle Beetles were also independent and eliminated the need for the equalizing torsion spring.

This new rear suspension layout would eventually become an option on later models. Beetles equipped with the automatic were identified with a "VW Automatic" badge on the engine lid and a matching decal in the rear window.

For , the only exterior change was the fuel filler flap no longer had a finger indentation due to a new interior-mounted fuel door release.

In other markets, manual transmission models retained a swing axle independent suspension which would continue until the end of German Beetle production.

In , a new "L" Luxus Package was introduced including, among other items, twin map pockets, dual rear ashtrays, full carpeting, a passenger-side visor vanity mirror, and rubber bumper moldings.

There were two Beetles for the first time in , the familiar standard Beetle and a new, larger version, different from the windscreen forward.

All Beetles received an engine upgrade: The ventilation system was improved with the original dash-top vents augmented by a second pair aimed directly at the driver and passenger.

For the first time the system was a flow-through design with crescent-shaped air exits fitted behind the rear quarter windows.

Airflow could be increased via an optional 2-speed fan. The standard Beetle was now badged as the VW ; when equipped with the engine, it was badged S, to avoided confusion with the Type 3, which wore VW badges.

The new suspension layout allowed the spare tire to be positioned flat under the trunk floor. Although the car had to be lengthened slightly to accomplish this, it allowed a reduction in turning radius.

To gain additional trunk volume, the under-dash panel [ clarification needed ] was lowered, allowing the fuel tank to be shifted rearward. From the windscreen back the big Beetle was identical to its smaller progenitor, except for having the also new semi-trailing arm rear suspension as standard equipment.

Both Beetles were available with or without the L Package. The convertible was now based on the body. While it lacked the front disc brakes that normally accompanied the larger motor, it was fitted with brake drums that were slightly larger than the standard Beetle.

With the Super Beetle being sold as the premium model in North America, the standard Beetle, while retaining the same cc engine, was stripped of many of its earlier features in order to reduce the selling price.

An engine compartment socket for the proprietary VW Diagnosis system was also introduced. The rear luggage area was fitted with a folding parcel shelf.

In the engine bay, the oil-bath air cleaner gave way to a dry element filter, and the generator was replaced with an alternator. The changes to the cowl and windshield resulted in slight redesign of the front hood.

The instrument panel, formerly shared with the standard Beetle, was all-new and incorporated a raised speedometer pod, rocker-style switches and side-window defrosters.

Front and rear deck lids were finished in matte black, as was all exterior trim with the exception of the chrome headlamp bezels.

Inside were corduroy and leatherette high-bolstered sport seats and a small diameter three-spoke steering wheel with padded leather rim and a small red VW logo [67] on the bottom spoke.

In some markets, the sport wheels in both 4. On the Super Beetle, the steering knuckle, and consequently the lower attachment point of the strut, was redesigned to improve handling and stability in the event of a tire blowout.

A limited-edition Big Beetle was introduced based on the LS. Available in unique metallic paint colors, the car featured styled-steel 5.

Both were finished in metallic gold and featured styled-steel 4. Inside were brown corduroy and leatherette seats, loop-pile carpet, and padded four-spoke deluxe steering wheel.

The Super Beetle Sun Bug included a sliding-steel sunroof. In , front turn indicators were moved from the top of the front fenders down into the bumper.

At the rear, the license plate light housing was now molded of plastic with a ribbed top surface. To comply with tightening emission standards, the cc engine in Japanese and North American markets received Bosch L-Jetronic fuel injection, rather than the D-Jetronic system formerly used in the VW Type 3 and Type 4.

The injected engine received a new muffler and in California a catalytic converter. This necessitated a bulge in the rear apron under the rear bumper and replaced the distinctive twin "pea shooter" tailpipes with a single offset pipe, making injected models identifiable at a glance.

The received rack and pinion steering. Mid-year, the Love Bug was introduced for North America: A price leader, the Love Bug retailed for less than a standard Beetle.

To make up for the loss in North American markets, the standard Beetle was upgraded, regaining some of the features that were removed in In addition, the 2-speed ventilation fan was included, previously available in North America only on the larger Beetle.

The automatic stickshift option was discontinued as well. This was the final model year for the Beetle sedan in North America. The convertible was offered in a "triple white" Champagne Edition in Alpine White with white top and interior with the padded deluxe steering wheel, tiger maple wood-grain dash trim and 4.

Approximately 1, Champagne Editions were produced. For , a new Champagne 2nd Edition convertible was launched, available in blue or red metallic paint with white leatherette interior.

Features included the 4. Approximately 1, were produced. In , VW offered an Epilogue Edition of the convertible in triple black with features similar to the Champagne Edition.

This would be the last year of convertible production worldwide as well as the final year for the Beetle in the USA and Canada. Hitler saw the car in at an auto show.

These small cars were designed according to the patents by Josef Ganz and featured transverse, two-stroke, two-cylinder engines mounted in front of the rear axle.

In , Tatra built the V prototype, which had an air-cooled flat-twin engine mounted at the rear. Both Hitler and Porsche were influenced by the Tatras.

It also featured a similar central structural tunnel found in the Beetle. Just before the start of the Second World War, Tatra had ten legal claims filed against VW for infringement of patents.

The vehicles were assembled in a former tram depot at Shelbourne Road in Ballsbridge. This is now the premises for Ballsbridge Motors which is still a Volkswagen dealer.

The first Volkswagen ever assembled outside Germany was built here. Jack Gilder had been fascinated by both the design and engineering of the Beetle when he came across one in Belgium during the war.

The Type 1 was introduced to Japan in , and was imported by Yanase dealerships in Japan. Its exterior dimensions and engine displacement were in compliance with Japanese Government regulations , which helped sales.

Several Japanese vehicles were introduced after the Beetle was sold in Japan, using an air-cooled engine and rear mounting of the engine, such as the Subaru , or an engine installed in the front, like the Honda N , the Suzuki Fronte , and the Mitsubishi Minica.

The last German made cars were assembled at Emden in , after which the Puebla, Mexico plant became the principal source of Beetle production.

Other countries produced Beetles from CKD complete knockdown kits: Brazilian assembly of the Beetle, where it is called " Fusca ", started in , with parts imported from Germany.

In production resumed and continued to The Brazilian version retained the —64 body style Europe and U. This body style was also produced in Mexico until The and s had the era taillights and headlights, fuel tank, but fitted with the up raised bumpers.

Brazilian CKD kits were shipped to Nigeria between and where Beetles were locally assembled. The Brazilian-produced versions have been sold in neighboring South American nations bordering Brazil, including Argentina, Uruguay, and Peru.

The Brazilian Type 1s have four different engines: In Brazil, the Type 1 never received electronic fuel injection, instead retaining carburetors one or two one-barrels throughout its entire life, although the carburetion differs from engines of different years and specification.

The production of the air-cooled engine finally ended in , after more than 60 years. It was last used in the Brazilian version of the VW Bus, called the "Kombi", and was replaced by a 1.

Volkswagen do Brasil engaged in some string pulling in the early sixties when a law requiring taxis to have four doors and five seats was being considered.

After proving that the average taxi fare only carried 1. Mexican production began in because of agreements with companies such as Chrysler in Mexico and the Studebaker-Packard Corporation which assembled cars imported in CKD form.

In , they began to be locally produced. These models have the larger windshield, rear window, door and quarter glass starting in ; and the rear window from to German built models was used on the Mexican models from to , when it was replaced with the larger rear window used on and later German built Beetles.

This version, after the mids, saw little change with the incorporation of electronic ignition in , an anti-theft alarm system in , a catalytic converter in as required by law , as well as electronic Digifant fuel injection , hydraulic valve lifters, and a spin-on oil filter in Starting in , the Mexican Beetle included front disc brakes, an alternator instead of a generator, and front automatic seat belts.

During the model year, the chrome moldings disappeared leaving body colored bumpers and black moldings instead on some models. These two versions were sold until mid Independent importers continued to supply several major countries, including Germany, France, and the UK until the end of production in Devoted fans of the car even discovered a way to circumvent US safety regulations by placing more recently manufactured Mexican Beetles on the floorpans of earlier, US-registered cars.

Some of the Mexican Beetles have been registered in the United States since the NHTSA amendment granting the year cutoff where it and its Brazilian counterpart including the T2C can be legally registered in any of the 50 states this means a or earlier Mexican Beetle as of can be registered under the current NHTSA year cutoff exemption.

The end of production in Mexico can be attributed primarily to Mexican political measures: The last Vocho taxis in Mexico City were retired at the end of In the late s consumers strongly preferred more modern cars such as the Mexican Chevy , the Nissan Tsuru , and the Volkswagen Pointer and Lupo.

However, demand for the Beetle was relatively high, especially in the s, with the workforce increasing at the start of the decade.

The price of the base model without even a radio was pegged with the official minimum wage, by an agreement between the company and the government.

Official importation of the Volkswagen Beetle into Australia began in , with local assembly operations commencing the following year. Volkswagen Australia was formed in , and by locally produced body panels were being used for the first time.

When the European Type One body received the larger windows for the model year, Volkswagen Australia decided not to update, but continued to produce the smaller-windowed bodies, with unique features to the Australian versions.

This was due to the limited size of the market and the costs involved in retooling. In , Volkswagen Australia released its own locally designed utilitarian version of the Type 1, the Volkswagen Country Buggy or Type The last Australian-assembled Beetle was produced in July with assembly of other models ending in February Several features from the Super Beetle were grafted onto the South African Beetle S, such as curved windshield, new dashboard, and larger taillights, while retaining the Beetle chassis and mechanicals.

The model was introduced to South Africa in ; it was marketed as the cheapest 1. In late , the sporty SP Beetle arrived — this version received bright red, yellow, or silver paint with black stripes, a front spoiler, wide tyres, and a more powerful engine with twin carburettors and a freer flowing exhaust.

Also new for were rubber bumper strips for all s, and the same taillights with backup lights were now fitted across the range.

The bigger-engined model was then phased out around August , leaving only the model in production. The Beetle has been modified for use in drag racing ; [95] its rearward RR layout weight distribution keeps the weight over the rear wheels, maximizing grip off the starting line.

To prevent this, "wheelie bars" were added. The Beetle is also used as the basis for the Formula Vee open-wheel racing category: In this configuration, FV would become one of the most popular entry-level motorsports classes of its time.

Volkswagen Beetle-style bodies are fitted to space frame racing chassis, and are used in the Uniroyal Fun Cup , which includes the longest continuous motor-race in the world, the 25 Hours of Spa.

The parents prepare the carcass and protect it from competitors and from early decomposition. After their eggs hatch, the parents keep the larvae clean of fungus and bacteria and help the larvae feed by regurgitating food for them.

Some dung beetles provide parental care, collecting herbivore dung and laying eggs within that food supply, an instance of mass provisioning.

Some species do not leave after this stage, but remain to safeguard their offspring. Most species of beetles do not display parental care behaviors after the eggs have been laid.

Eusociality involves cooperative brood care including brood care of offspring from other individuals , overlapping generations within a colony of adults, and a division of labour into reproductive and non-reproductive groups.

It is one of more than species of wood-boring Ambrosia beetles which distribute the spores of ambrosia fungi. Beetles are able to exploit a wide diversity of food sources available in their many habitats.

Some are omnivores , eating both plants and animals. Other beetles are highly specialized in their diet. Many species of leaf beetles, longhorn beetles, and weevils are very host-specific, feeding on only a single species of plant.

Ground beetles and rove beetles Staphylinidae , among others, are primarily carnivorous and catch and consume many other arthropods and small prey, such as earthworms and snails.

While most predatory beetles are generalists, a few species have more specific prey requirements or preferences.

Decaying organic matter is a primary diet for many species. This can range from dung , which is consumed by coprophagous species such as certain scarab beetles in the Scarabaeidae , to dead animals, which are eaten by necrophagous species such as the carrion beetles , Silphidae.

Some beetles found in dung and carrion are in fact predatory. These include members of the Histeridae and Silphidae , preying on the larvae of coprophagous and necrophagous insects.

Some beetles have special mycangia , structures for the transport of fungal spores. Beetles, both adults and larvae, are the prey of many animal predators including mammals from bats to rodents , birds , lizards , amphibians , fishes , dragonflies , robberflies , reduviid bugs , ants , other beetles, and spiders.

These include camouflage and mimicry against predators that hunt by sight, toxicity, and defensive behaviour.

Camouflage is common and widespread among beetle families, especially those that feed on wood or vegetation, such as leaf beetles Chrysomelidae, which are often green and weevils.

In some species, sculpturing or various coloured scales or hairs cause beetles such as the avocado weevil Heilipus apiatus to resemble bird dung or other inedible objects.

Some longhorn beetles Cerambycidae are effective Batesian mimics of wasps. Beetles may combine coloration with behavioural mimicry, acting like the wasps they already closely resemble.

Many other beetles, including ladybirds , blister beetles , and lycid beetles secrete distasteful or toxic substances to make them unpalatable or poisonous, and are often aposematic , where bright or contrasting coloration warn off predators; many beetles and other insects mimic these chemically protected species.

Chemical defense is important in some species, usually being advertised by bright aposematic colours. Some Tenebrionidae use their posture for releasing noxious chemicals to warn off predators.

Chemical defences may serve purposes other than just protection from vertebrates, such as protection from a wide range of microbes.

Some species sequester chemicals from the plants they feed on, incorporating them into their own defenses. Other species have special glands to produce deterrent chemicals.

The defensive glands of carabid ground beetles produce a variety of hydrocarbons , aldehydes , phenols , quinones , esters , and acids released from an opening at the end of the abdomen.

African carabid beetles for example, Anthia and Thermophilum — Thermophilum is sometimes included within Anthia employ the same chemicals as ants: The gland is made of two containing chambers, one for hydroquinones and hydrogen peroxide , the other holding hydrogen peroxide and catalase enzymes.

The oxygen propels the noxious chemical spray as a jet that can be aimed accurately at predators. Large ground-dwelling beetles such as Carabidae , the rhinoceros beetle and the longhorn beetles defend themselves using strong mandibles , or heavily sclerotised armored spines or horns to deter or fight off predators.

Some combine it with thanatosis , in which they close up their appendages and "play dead". A few species of beetles are ectoparasitic on mammals.

One such species, Platypsyllus castoris , parasitises beavers Castor spp. This beetle lives as a parasite both as a larva and as an adult, feeding on epidermal tissue and possibly on skin secretions and wound exudates.

They are wingless and eyeless, as are many other ectoparasites. Beetle-pollinated flowers are usually large, greenish or off-white in color, and heavily scented.

Scents may be spicy, fruity, or similar to decaying organic material. Beetles were most likely the first insects to pollinate flowers. Most beetle-pollinated flowers are flattened or dish-shaped, with pollen easily accessible, although they may include traps to keep the beetle longer.

The beetle families that habitually pollinate flowers are the Buprestidae, Cantharidae, Carambycidae, Cleridae, Dermestidae, Lycidae, Melyridae, Mordellidae, Nitidulidae and Scarabeidae.

Mutualism is well known in a few beetles, such as the ambrosia beetle , which partners with fungi to digest the wood of dead trees.

The beetles excavate tunnels in dead trees in which they cultivate fungal gardens, their sole source of nutrition.

After landing on a suitable tree, an ambrosia beetle excavates a tunnel in which it releases spores of its fungal symbiont.

The beetles cannot eat the wood due to toxins, and uses its relationship with fungi to help overcome the defenses of its host tree in order to provide nutrition for their larvae.

Adult diapause is the most common form of diapause in Coleoptera. To endure the period without food often lasting many months adults prepare by accumulating reserves of lipids, glycogen, proteins and other substances needed for resistance to future hazardous changes of environmental conditions.

This diapause is induced by signals heralding the arrival of the unfavourable season; usually the cue is photoperiodic.

Short decreasing day length serves as a signal of approaching winter and induces winter diapause hibernation. This loss of body fat was a gradual process, occurring in combination with dehydration.

All insects are poikilothermic , [] so the ability of a few beetles to live in extreme environments depends on their resilience to unusually high or low temperatures.

The low temperatures experienced by Cucujus clavipes can be survived through their deliberate dehydration in conjunction with the antifreeze proteins.

This concentrates the antifreezes several fold. Conversely, desert dwelling beetles are adapted to tolerate high temperatures.

These beetles also exhibits behavioural adaptions to tolerate the heat: The fogstand beetle of the Namib Desert , Stenocara gracilipes , is able to collect water from fog , as its elytra have a textured surface combining hydrophilic water-loving bumps and waxy, hydrophobic troughs.

The beetle faces the early morning breeze, holding up its abdomen; droplets condense on the elytra and run along ridges towards their mouthparts.

Similar adaptations are found in several other Namib desert beetles such as Onymacris unguicularis. Some terrestrial beetles that exploit shoreline and floodplain habitats have physiological adaptations for surviving floods.

In the event of flooding, adult beetles may be mobile enough to move away from flooding, but larvae and pupa often cannot.

Adults of Cicindela togata are unable to survive immersion in water, but larvae are able to survive a prolonged period, up to 6 days, of anoxia during floods.

Anoxia tolerance in the larvae may have been sustained by switching to anaerobic metabolic pathways or by reducing metabolic rate.

Many beetle species undertake annual mass movements which are termed as migrations. These include the pollen beetle Meligethes aeneus [] and many species of coccinellids.

Several species of dung beetle, especially the sacred scarab, Scarabaeus sacer , were revered in Ancient Egypt. The scarab was of prime significance in the funerary cult of ancient Egypt.

The best-known of these are the Judean LMLK seals , where eight of 21 designs contained scarab beetles, which were used exclusively to stamp impressions on storage jars during the reign of Hezekiah.

Pliny the Elder discusses beetles in his Natural History , [] describing the stag beetle: To these insects a sting has been denied by Nature; but in one large kind we find horns of a remarkable length, two-pronged at the extremities, and forming pincers, which the animal closes when it is its intention to bite.

He is black, long and has hard wings like a great dung beetle". Many feed on economically important plants and stored plant products, including trees, cereals, tobacco, and dried fruits.

The boll weevil crossed the Rio Grande near Brownsville , Texas, to enter the United States from Mexico around , [] and had reached southeastern Alabama by It remains the most destructive cotton pest in North America.

The bark beetle, elm leaf beetle and the Asian longhorned beetle Anoplophora glabripennis [] are among the species that attack elm trees.

Bark beetles Scolytidae carry Dutch elm disease as they move from infected breeding sites to healthy trees. The disease has devastated elm trees across Europe and North America.

Some species of beetle have evolved immunity to insecticides. For example, the Colorado potato beetle , Leptinotarsa decemlineata , is a destructive pest of potato plants.

Its hosts include other members of the Solanaceae , such as nightshade , tomato , eggplant and capsicum , as well as the potato. Different populations have between them developed resistance to all major classes of insecticide.

The death watch beetle , Xestobium rufovillosum Ptinidae , is a serious pest of older wooden buildings in Europe. It attacks hardwoods such as oak and chestnut , always where some fungal decay has taken or is taking place.

The actual introduction of the pest into buildings is thought to take place at the time of construction. Other pests include the coconut hispine beetle, Brontispa longissima , which feeds on young leaves , seedlings and mature coconut trees, causing serious economic damage in the Philippines.

Beetles can be beneficial to human economics by controlling the populations of pests. The larvae and adults of some species of lady beetles Coccinellidae feed on aphids that are pests.

Other lady beetles feed on scale insects , whitefly and mealybugs. For example, the genus Zygogramma is native to North America but has been used to control Parthenium hysterophorus in India and Ambrosia artemisiifolia in Russia.

Dung beetles Scarabidae have been successfully used to reduce the populations of pestilent flies, such as Musca vetustissima and Haematobia exigua which are serious pests of cattle in Australia.

The Dermestidae are often used in taxidermy and in the preparation of scientific specimens, to clean soft tissue from bones. Beetles are the most widely eaten insects, with about species used as food, usually at the larval stage.

Due to their habitat specificity, many species of beetles have been suggested as suitable as indicators, their presence, numbers, or absence providing a measure of habitat quality.

Predatory beetles such as the tiger beetles Cicindelidae have found scientific use as an indicator taxon for measuring regional patterns of biodiversity.

They are suitable for this as their taxonomy is stable; their life history is well described; they are large and simple to observe when visiting a site; they occur around the world in many habitats, with species specialised to particular habitats; and their occurrence by species accurately indicates other species, both vertebrate and invertebrate.

Many beetles have beautiful and durable elytra that have been used as material in arts, with beetlewing the best example.

Whole beetles, either as-is or encased in clear plastic, are made into objects ranging from cheap souvenirs such as key chains to expensive fine-art jewellery.

In parts of Mexico, beetles of the genus Zopherus are made into living brooches by attaching costume jewelry and golden chains, which is made possible by the incredibly hard elytra and sedentary habits of the genus.

Fighting beetles are used for entertainment and gambling. This sport exploits the territorial behavior and mating competition of certain species of large beetles.

In the Chiang Mai district of northern Thailand, male Xylotrupes rhinoceros beetles are caught in the wild and trained for fighting.

Females are held inside a log to stimulate the fighting males with their pheromones. Beetles are sometimes used as instruments: Some species of beetle are kept as pets , for example diving beetles Dytiscidae may be kept in a domestic fresh water tank.

In Japan the practice of keeping horned rhinoceros beetles Dynastinae and stag beetles Lucanidae is particularly popular amongst young boys.

Beetle collecting became extremely popular in the Victorian era. Several coleopteran adaptations have attracted interest in biomimetics with possible commercial applications.

Living beetles have been used as cyborgs. A Defense Advanced Research Projects Agency funded project implanted electrodes into Mecynorhina torquata beetles, allowing them to be remotely controlled via a radio receiver held on its back, as proof-of-concept for surveillance work.

Many species of beetles have very specific habitats and long life cycles that make them vulnerable. Some species are highly threatened while others are already feared extinct.

In Japan the Genji firefly, Luciola cruciata , is extremely popular, and in South Africa the Addo elephant dung beetle offers promise for broadening ecotourism beyond the big five tourist mammal species.

Popular dislike of pest beetles, too, can be turned into public interest in insects, as can unusual ecological adaptations of species like the fairy shrimp hunting beetle, Cicinis bruchi.

From Wikipedia, the free encyclopedia. This article is about the insect. For the car, see Volkswagen Beetle. For other uses, see Beetle disambiguation.

Insect thermoregulation and Insect winter ecology. Beetlewing and Live insect jewelry. Retrieved 20 February Species Diversity in Space and Time.

An Outline of Entomology 5 ed. Book of Insect Records. Archived from the original on July 18, New record and remeasuring of Scydosella musawasensis Hall, Coleoptera, Ptiliidae , the smallest known free-living insect".

The beetle fauna of Germany. Retrieved March 16, Evolution of the Insects. Triassic Curculionoidea have the same status as Triassic Chrysoleloidea: Journal of the Kansas Entomological Society.

Archived from the original PDF on July 18, Archived from the original PDF on November 11, Archived from the original PDF on March 31, Archived from the original PDF on July 4, Annalen des Naturhistorischen Museums in Wien.

Archived from the original PDF on The origin of coprophagy ". Journal of Natural History. Proceedings of the Royal Society B: Evolution, Biogeography, and Ecology".

Annual Review of Ecology and Systematics. Tree of Life web project. Retrieved 27 February Handbook of Zoology, Band 4: Morphology, Classification and Keys.

A new beetle family to the fauna of Serbia". Bulletin of the Natural History Museum 6: Carabidae and Trachypachidae ". Tree of Life Web Project.

Retrieved March 18, Retrieved 15 March Journal of Asia-Pacific Entomology. A Guide to the Beetles of Australia.

American Beetles, Volume I: Archostemata, Myxophaga, Adephaga, Polyphaga: Head, Thorax, Abdomen, and Genitalia Insects ".

Journal of the Royal Society Interface. University of Minnesota Extension. Insects, their structure and life. Entomologische Berichten in Dutch.

The Journal of Experimental Biology. Adaptation and Environment 5th ed. Structure and Physiology of the Circulatory System".

Comprehensive Insect Physiology, Biochemistry and Pharmacology. Integument, Respiration and Circulation. Form, function, and diversity. Turkish Journal of Zoology.

The Biology of the Coleoptera. Medical and Veterinary Entomology 2nd ed. Retrieved 17 March Insights from Vertical-Looking Entomological Radar".

Encyclopedia of Entomology 2 ed. American Beetles, Volume 1. Retrieved March 13, Journal of the New York Entomological Society. The Australian beetle that behaves like a bee".

Beetle Biology And Ecology. Beetles Coleoptera and Coleopterologist. Jahangirnagar University Journal of Biological Science.

Adaptive Coloration in Animals. Gaps in our knowledge of an invasive species". North Carolina State University. Taxonomy, Ecology and Pathology.

Adult Diapause in Coleoptera". A Journal of Entomology. An Outline of Entomology. Journal of Experimental Biology. Surviving in hostile habitats".

The yellow mealworm beetle, Tenebrio molitor, contains a family of small Cys-rich and Thr-rich thermal hysteresis proteins that depress the hemolymph freezing point below the melting point by as much as 5.

Thermal hysteresis protein expression was evaluated throughout development and after exposure to altered environmental conditions. Journal of Biological Chemistry.

Annals of the Entomological Society of America. Wyatt; Stanley, David W. The American Midland Naturalist. The Biology of Life on the Move. Retrieved 10 February Retrieved July 19, The New Archaeological Excavations at Lachish — Institute of Archaeology of Tel Aviv University.

University of Chicago Press. Insects and other Invertebrates in Classical Antiquity. The Metamorphoses of Antoninus Liberalis: A translation with a commentary.

London and New York, Routledge. Economic impacts of the boll weevil. Archived from the original on May 12, Canadian Journal of Plant Pathology.

American Journal of Potato Research. Why our next terrorist attack could come on six legs". Pest Management and Phytosanitary Trade Barriers. Conservation and collective care.

Archived from the original on July 10, Retrieved July 17, Chrysomelidae in the field". Archived from the original on April 19, Retrieved June 24, Retrieved June 17, Agriculture, Ecosystems and Environment.

Retrieved 14 March Coleoptera, Chrysomelidae in biological control of ragweeds Ambrosia artemisiifolia L. Entomologicheskoe Obozrenije in Russian.

Australian Journal of Entomology. Scarabaeidae can improve soil hydrological properties". In Huffman, Jane E.

Developments in Forensic Science. Preparation protocols for skeletons and fossil vertebrates under the scanning electron microscopy". Retrieved 26 January Journal of Ethnobiology and Ethnomedicine.

Indicator Taxon for Biodiversity and Conservation Studies". A case study of dung beetles in a savanna ecosystem". Journal of Applied Ecology.

Arnett; Michael Charles Thomas. Volume 2 of American Beetles. A brief review" PDF. Entomology in modern Japan" PDF.

Representations of Animals in Victorian Literature and Culture. The land of the orang—utan, and the bird of paradise. A narrative of travel, with sketches of man and nature 1 ed.

Explanations of carabid beetle Coleoptera, Carabidae declines in Europe". An Inordinate Fondness for Beetles. University of California Press.

Encyclopedia of Insects 2nd ed. Thysanura Zygentoma silverfish, firebrats. Plecoptera stoneflies Dermaptera earwigs Embioptera webspinners Phasmatodea stick and leaf insects Notoptera ice-crawlers, gladiators Orthoptera crickets, wetas, grasshoppers, locusts Zoraptera angel insects.

Blattodea cockroaches, termites Mantodea mantises. Psocodea barklice, lice Thysanoptera thrips Hemiptera cicadas, aphids, true bugs.

Hymenoptera sawflies, wasps, ants, bees. Strepsiptera twisted-winged parasites Coleoptera beetles. Raphidioptera snakeflies Megaloptera alderflies, dobsonflies, fishflies Neuroptera net-winged insects: Trichoptera caddisflies Lepidoptera moths, butterflies.

Four most speciose orders are marked in bold Italic are paraphyletic groups Based on Sasaki et al. Extinct incertae sedis families and genera are marked in italic.

Crowsoniellidae Crowsoniella relicta Cupedidae reticulated beetles Jurodidae Sikhotealinia zhiltzovae Micromalthidae telephone-pole beetle Ommatidae.

Amphizoidae trout-stream beetles Aspidytidae Carabidae ground beetles Dytiscidae predaceous diving beetles Gyrinidae whirligig beetles Haliplidae crawling water beetles Hygrobiidae Meruidae Meru phyllisae Noteridae burrowing water beetles Rhysodidae wrinkled bark beetles Trachypachidae false ground beetles.

Hydroscaphidae skiff beetles Lepiceridae Sphaeriusidae Torridincolidae. Derodontidae tooth-necked fungus beetles. Cerambycidae longhorn beetles Chrysomelidae leaf beetles Disteniidae Megalopodidae Orsodacnidae Oxypeltidae Vesperidae.

Acanthocnemidae Acanthocnemus nigricans Chaetosomatidae Cleridae checkered beetles Mauroniscidae Melyridae soft-wing flower beetles Metaxinidae Phloiophilidae Phloiophilus edwardsi Phycosecidae Prionoceridae Thanerocleridae Trogossitidae bark-gnawing beetles.

Anthribidae fungus weevils Attelabidae leaf-rolling weevils Belidae primitive weevils Brentidae straight snout weevils, New York weevil Caridae Curculionidae true weevils, bark beetles, ambrosia beetles Nemonychidae pine flower weevils.

Aderidae ant-like leaf beetles Anthicidae ant-like flower beetles Archeocrypticidae cryptic fungus beetles Boridae conifer bark beetles Chalcodryidae Ciidae minute tree-fungus beetles Melandryidae false darkling beetles Meloidae blister beetles Mordellidae tumbling flower beetles Mycetophagidae hairy fungus beetles Mycteridae palm and flower beetles Oedemeridae false blister beetle Perimylopidae, or Promecheilidae Prostomidae jugular-horned beetles Pterogeniidae Pyrochroidae fire-coloured beetles Pythidae dead log bark beetles Ripiphoridae wedge-shaped beetles Salpingidae narrow-waisted bark beetles Scraptiidae false flower beetles Stenotrachelidae false longhorn beetles Synchroidae synchroa bark beetles Tenebrionidae darkling beetles Tetratomidae polypore fungus beetles Trachelostenidae Trictenotomidae Ulodidae Zopheridae ironclad beetles, cylindrical bark beetles.

Buprestidae jewel beetles, or metallic wood-boring beetles Schizopodidae. Byrrhidae pill beetles Callirhipidae cedar beetles Chelonariidae turtle beetles Cneoglossidae Dryopidae long-toed water beetles Elmidae riffle beetles Eulichadidae forest stream beetles Heteroceridae variegated mud-loving beetles Limnichidae minute mud beetles Lutrochidae travertine beetles Psephenidae water-penny beetles Ptilodactylidae.

Dascillidae soft bodied plant beetles Rhipiceridae cicada beetle, cicada parasite beetles. Anischiidae Artematopodidae soft-bodied plant beetles Brachypsectridae Texas beetles Cantharidae soldier beetles Cerophytidae rare click beetles Elateridae click beetles Eucnemidae false click beetles Lampyridae fireflies Lycidae net-winged beetles Omalisidae Omethidae false fireflies Phengodidae glowworm beetles, long-lipped beetles Podabrocephalidae Rhagophthalmidae Rhinorhipidae Rhinorhipus tamborinensis Throscidae false metallic wood-boring beetles.

Clambidae Decliniidae Declinia relicta Eucinetidae plate-thigh beetles Scirtidae.

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Dermestids produce esters , and species of Elateridae produce fatty acid-derived aldehydes and acetates. Light production is highly efficient, by oxidation of luciferin catalyzed by enzymes luciferases in the presence of adenosine triphosphate ATP and oxygen, producing oxyluciferin , carbon dioxide, and light.

Tympanal organs or hearing organs consist of a membrane tympanum stretched across a frame backed by an air sac and associated sensory neurons, are found in two families.

Both families are sensitive to ultrasonic frequencies, with strong evidence indicating they function to detect the presence of bats by their ultrasonic echolocation.

Beetles are members of the superorder Endopterygota , and accordingly most of them undergo complete metamorphosis. The typical form of metamorphosis in beetles passes through four main stages: The larvae are commonly called grubs and the pupa sometimes is called the chrysalis.

In some species, the pupa may be enclosed in a cocoon constructed by the larva towards the end of its final instar.

Some beetles, such as typical members of the families Meloidae and Rhipiphoridae , go further, undergoing hypermetamorphosis in which the first instar takes the form of a triungulin.

Some beetles have intricate mating behaviour. Pheromone communication is often important in locating a mate. Different species use different pheromones.

Scarab beetles such as the Rutelinae use pheromones derived from fatty acid synthesis , while other scarabs such as the Melolonthinae use amino acids and terpenoids.

Another way beetles find mates is seen in the fireflies Lampyridae which are bioluminescent , with abdominal light-producing organs. The males and females engage in a complex dialogue before mating; each species has a unique combination of flight patterns, duration, composition, and intensity of the light produced.

Before mating, males and females may stridulate, or vibrate the objects they are on. In the Meloidae, the male climbs onto the dorsum of the female and strokes his antennae on her head, palps, and antennae.

In Eupompha , the male draws his antennae along his longitudinal vertex. They may not mate at all if they do not perform the precopulatory ritual.

Competition can play a part in the mating rituals of species such as burying beetles Nicrophorus , the insects fighting to determine which can mate.

Many male beetles are territorial and fiercely defend their territories from intruding males. In such species, the male often has horns on the head or thorax, making its body length greater than that of a female.

Copulation is generally quick, but in some cases lasts for several hours. During copulation, sperm cells are transferred to the female to fertilize the egg.

Essentially all beetles lay eggs, though some myrmecophilous Aleocharinae and some Chrysomelinae which live in mountains or the subarctic are ovoviviparous , laying eggs which hatch almost immediately.

Beetle eggs generally have smooth surfaces and are soft, though the Cupedidae have hard eggs. Eggs vary widely between species: A female may lay from several dozen to several thousand eggs during her lifetime, depending on the extent of parental care.

This ranges from the simple laying of eggs under a leaf, to the parental care provided by scarab beetles , which house, feed and protect their young.

The Attelabidae roll leaves and lay their eggs inside the roll for protection. The larva is usually the principal feeding stage of the beetle life cycle.

Larvae tend to feed voraciously once they emerge from their eggs. Some feed externally on plants, such as those of certain leaf beetles, while others feed within their food sources.

Examples of internal feeders are most Buprestidae and longhorn beetles. The larvae of many beetle families are predatory like the adults ground beetles, ladybirds, rove beetles.

The larval period varies between species, but can be as long as several years. The larvae of skin beetles undergo a degree of reversed development when starved, and later grow back to the previously attained level of maturity.

The cycle can be repeated many times see Biological immortality. Beetle larvae can be differentiated from other insect larvae by their hardened, often darkened heads, the presence of chewing mouthparts, and spiracles along the sides of their bodies.

Like adult beetles, the larvae are varied in appearance, particularly between beetle families. Beetles with somewhat flattened, highly mobile larvae include the ground beetles and rove beetles; their larvae are described as campodeiform.

Some beetle larvae resemble hardened worms with dark head capsules and minute legs. These are elateriform larvae, and are found in the click beetle Elateridae and darkling beetle Tenebrionidae families.

Some elateriform larvae of click beetles are known as wireworms. Beetles in the Scarabaeoidea have short, thick larvae described as scarabaeiform, more commonly known as grubs.

All beetle larvae go through several instars , which are the developmental stages between each moult. In many species, the larvae simply increase in size with each successive instar as more food is consumed.

In some cases, however, more dramatic changes occur. Among certain beetle families or genera, particularly those that exhibit parasitic lifestyles, the first instar the planidium is highly mobile to search out a host, while the following instars are more sedentary and remain on or within their host.

This is known as hypermetamorphosis ; it occurs in the Meloidae , Micromalthidae , and Ripiphoridae. Its first stage, the triungulin , has longer legs to go in search of the eggs of grasshoppers.

After feeding for a week it moults to the second stage, called the caraboid stage, which resembles the larva of a carabid beetle. In another week it moults and assumes the appearance of a scarabaeid larva — the scarabaeidoid stage.

Its penultimate larval stage is the pseudo-pupa or the coarcate larva, which will overwinter and pupate until the next spring. The larval period can vary widely.

A fungus feeding staphylinid Phanerota fasciata undergoes three moults in 3. Leiodidae completes its larval stage in the fruiting body of slime mold in 2 days and possibly represents the fastest growing beetles.

Dermestid beetles, Trogoderma inclusum can remain in an extended larval state under unfavourable conditions, even reducing their size between moults.

A larva is reported to have survived for 3. As with all endopterygotes, beetle larvae pupate, and from these pupae emerge fully formed, sexually mature adult beetles, or imagos.

Pupae never have mandibles they are adecticous. In most pupae, the appendages are not attached to the body and are said to be exarate ; in a few beetles Staphylinidae, Ptiliidae etc.

Adults have extremely variable lifespans, from weeks to years, depending on the species. It is believed that when furniture or house timbers are infested by beetle larvae, the timber already contained the larvae when it was first sawn up.

A birch bookcase 40 years old released adult Eburia quadrigeminata Cerambycidae , while Buprestis aurulenta and other Buprestidae have been documented as emerging as much as 51 years after manufacture of wooden items.

The elytra allow beetles to both fly and move through confined spaces, doing so by folding the delicate wings under the elytra while not flying, and folding their wings out just before take off.

The unfolding and folding of the wings is operated by muscles attached to the wing base; as long as the tension on the radial and cubital veins remains, the wings remain straight.

In some day-flying species for example, Buprestidae , Scarabaeidae , flight does not include large amounts of lifting of the elytra, having the metathorac wings extended under the lateral elytra margins.

Many rove beetles have greatly reduced elytra, and while they are capable of flight, they most often move on the ground: Diving beetles Dytiscidae hold air between the abdomen and the elytra when diving.

Hydrophilidae have hairs on their under surface that retain a layer of air against their bodies. Adult crawling water beetles use both their elytra and their hind coxae the basal segment of the back legs in air retention, while whirligig beetles simply carry an air bubble down with them whenever they dive.

Beetles have a variety of ways to communicate, including the use of pheromones. The mountain pine beetle emits a pheromone to attract other beetles to a tree.

The mass of beetles are able to overcome the chemical defenses of the tree. This species can stridulate to communicate [88] , but others may use sound to defend themselves when attacked [89].

Parental care is found in a few species of beetle, perhaps for protection against adverse conditions and predators.

Burying beetles are attentive parents, and participate in cooperative care and feeding of their offspring. Both parents work to bury small animal carcass to serve as a food resource for their young and build a brood chamber around it.

The parents prepare the carcass and protect it from competitors and from early decomposition. After their eggs hatch, the parents keep the larvae clean of fungus and bacteria and help the larvae feed by regurgitating food for them.

Some dung beetles provide parental care, collecting herbivore dung and laying eggs within that food supply, an instance of mass provisioning.

Some species do not leave after this stage, but remain to safeguard their offspring. Most species of beetles do not display parental care behaviors after the eggs have been laid.

Eusociality involves cooperative brood care including brood care of offspring from other individuals , overlapping generations within a colony of adults, and a division of labour into reproductive and non-reproductive groups.

It is one of more than species of wood-boring Ambrosia beetles which distribute the spores of ambrosia fungi. Beetles are able to exploit a wide diversity of food sources available in their many habitats.

Some are omnivores , eating both plants and animals. Other beetles are highly specialized in their diet. Many species of leaf beetles, longhorn beetles, and weevils are very host-specific, feeding on only a single species of plant.

Ground beetles and rove beetles Staphylinidae , among others, are primarily carnivorous and catch and consume many other arthropods and small prey, such as earthworms and snails.

While most predatory beetles are generalists, a few species have more specific prey requirements or preferences.

Decaying organic matter is a primary diet for many species. This can range from dung , which is consumed by coprophagous species such as certain scarab beetles in the Scarabaeidae , to dead animals, which are eaten by necrophagous species such as the carrion beetles , Silphidae.

Some beetles found in dung and carrion are in fact predatory. These include members of the Histeridae and Silphidae , preying on the larvae of coprophagous and necrophagous insects.

Some beetles have special mycangia , structures for the transport of fungal spores. Beetles, both adults and larvae, are the prey of many animal predators including mammals from bats to rodents , birds , lizards , amphibians , fishes , dragonflies , robberflies , reduviid bugs , ants , other beetles, and spiders.

These include camouflage and mimicry against predators that hunt by sight, toxicity, and defensive behaviour. Camouflage is common and widespread among beetle families, especially those that feed on wood or vegetation, such as leaf beetles Chrysomelidae, which are often green and weevils.

In some species, sculpturing or various coloured scales or hairs cause beetles such as the avocado weevil Heilipus apiatus to resemble bird dung or other inedible objects.

Some longhorn beetles Cerambycidae are effective Batesian mimics of wasps. Beetles may combine coloration with behavioural mimicry, acting like the wasps they already closely resemble.

Many other beetles, including ladybirds , blister beetles , and lycid beetles secrete distasteful or toxic substances to make them unpalatable or poisonous, and are often aposematic , where bright or contrasting coloration warn off predators; many beetles and other insects mimic these chemically protected species.

Chemical defense is important in some species, usually being advertised by bright aposematic colours. Some Tenebrionidae use their posture for releasing noxious chemicals to warn off predators.

Chemical defences may serve purposes other than just protection from vertebrates, such as protection from a wide range of microbes. Some species sequester chemicals from the plants they feed on, incorporating them into their own defenses.

Other species have special glands to produce deterrent chemicals. The defensive glands of carabid ground beetles produce a variety of hydrocarbons , aldehydes , phenols , quinones , esters , and acids released from an opening at the end of the abdomen.

African carabid beetles for example, Anthia and Thermophilum — Thermophilum is sometimes included within Anthia employ the same chemicals as ants: The gland is made of two containing chambers, one for hydroquinones and hydrogen peroxide , the other holding hydrogen peroxide and catalase enzymes.

The oxygen propels the noxious chemical spray as a jet that can be aimed accurately at predators. Large ground-dwelling beetles such as Carabidae , the rhinoceros beetle and the longhorn beetles defend themselves using strong mandibles , or heavily sclerotised armored spines or horns to deter or fight off predators.

Some combine it with thanatosis , in which they close up their appendages and "play dead". A few species of beetles are ectoparasitic on mammals.

One such species, Platypsyllus castoris , parasitises beavers Castor spp. This beetle lives as a parasite both as a larva and as an adult, feeding on epidermal tissue and possibly on skin secretions and wound exudates.

They are wingless and eyeless, as are many other ectoparasites. Beetle-pollinated flowers are usually large, greenish or off-white in color, and heavily scented.

Scents may be spicy, fruity, or similar to decaying organic material. Beetles were most likely the first insects to pollinate flowers. Most beetle-pollinated flowers are flattened or dish-shaped, with pollen easily accessible, although they may include traps to keep the beetle longer.

The beetle families that habitually pollinate flowers are the Buprestidae, Cantharidae, Carambycidae, Cleridae, Dermestidae, Lycidae, Melyridae, Mordellidae, Nitidulidae and Scarabeidae.

Mutualism is well known in a few beetles, such as the ambrosia beetle , which partners with fungi to digest the wood of dead trees.

The beetles excavate tunnels in dead trees in which they cultivate fungal gardens, their sole source of nutrition. After landing on a suitable tree, an ambrosia beetle excavates a tunnel in which it releases spores of its fungal symbiont.

The beetles cannot eat the wood due to toxins, and uses its relationship with fungi to help overcome the defenses of its host tree in order to provide nutrition for their larvae.

Adult diapause is the most common form of diapause in Coleoptera. To endure the period without food often lasting many months adults prepare by accumulating reserves of lipids, glycogen, proteins and other substances needed for resistance to future hazardous changes of environmental conditions.

This diapause is induced by signals heralding the arrival of the unfavourable season; usually the cue is photoperiodic. Short decreasing day length serves as a signal of approaching winter and induces winter diapause hibernation.

This loss of body fat was a gradual process, occurring in combination with dehydration. All insects are poikilothermic , [] so the ability of a few beetles to live in extreme environments depends on their resilience to unusually high or low temperatures.

The low temperatures experienced by Cucujus clavipes can be survived through their deliberate dehydration in conjunction with the antifreeze proteins.

This concentrates the antifreezes several fold. Conversely, desert dwelling beetles are adapted to tolerate high temperatures. These beetles also exhibits behavioural adaptions to tolerate the heat: The fogstand beetle of the Namib Desert , Stenocara gracilipes , is able to collect water from fog , as its elytra have a textured surface combining hydrophilic water-loving bumps and waxy, hydrophobic troughs.

The beetle faces the early morning breeze, holding up its abdomen; droplets condense on the elytra and run along ridges towards their mouthparts.

Similar adaptations are found in several other Namib desert beetles such as Onymacris unguicularis. Some terrestrial beetles that exploit shoreline and floodplain habitats have physiological adaptations for surviving floods.

In the event of flooding, adult beetles may be mobile enough to move away from flooding, but larvae and pupa often cannot. Adults of Cicindela togata are unable to survive immersion in water, but larvae are able to survive a prolonged period, up to 6 days, of anoxia during floods.

Anoxia tolerance in the larvae may have been sustained by switching to anaerobic metabolic pathways or by reducing metabolic rate.

Many beetle species undertake annual mass movements which are termed as migrations. These include the pollen beetle Meligethes aeneus [] and many species of coccinellids.

Several species of dung beetle, especially the sacred scarab, Scarabaeus sacer , were revered in Ancient Egypt.

The scarab was of prime significance in the funerary cult of ancient Egypt. The best-known of these are the Judean LMLK seals , where eight of 21 designs contained scarab beetles, which were used exclusively to stamp impressions on storage jars during the reign of Hezekiah.

Pliny the Elder discusses beetles in his Natural History , [] describing the stag beetle: To these insects a sting has been denied by Nature; but in one large kind we find horns of a remarkable length, two-pronged at the extremities, and forming pincers, which the animal closes when it is its intention to bite.

He is black, long and has hard wings like a great dung beetle". Many feed on economically important plants and stored plant products, including trees, cereals, tobacco, and dried fruits.

The boll weevil crossed the Rio Grande near Brownsville , Texas, to enter the United States from Mexico around , [] and had reached southeastern Alabama by It remains the most destructive cotton pest in North America.

The bark beetle, elm leaf beetle and the Asian longhorned beetle Anoplophora glabripennis [] are among the species that attack elm trees.

Bark beetles Scolytidae carry Dutch elm disease as they move from infected breeding sites to healthy trees. The disease has devastated elm trees across Europe and North America.

Some species of beetle have evolved immunity to insecticides. For example, the Colorado potato beetle , Leptinotarsa decemlineata , is a destructive pest of potato plants.

Its hosts include other members of the Solanaceae , such as nightshade , tomato , eggplant and capsicum , as well as the potato.

Different populations have between them developed resistance to all major classes of insecticide. The death watch beetle , Xestobium rufovillosum Ptinidae , is a serious pest of older wooden buildings in Europe.

It attacks hardwoods such as oak and chestnut , always where some fungal decay has taken or is taking place. The actual introduction of the pest into buildings is thought to take place at the time of construction.

Other pests include the coconut hispine beetle, Brontispa longissima , which feeds on young leaves , seedlings and mature coconut trees, causing serious economic damage in the Philippines.

Beetles can be beneficial to human economics by controlling the populations of pests. The larvae and adults of some species of lady beetles Coccinellidae feed on aphids that are pests.

Other lady beetles feed on scale insects , whitefly and mealybugs. For example, the genus Zygogramma is native to North America but has been used to control Parthenium hysterophorus in India and Ambrosia artemisiifolia in Russia.

Dung beetles Scarabidae have been successfully used to reduce the populations of pestilent flies, such as Musca vetustissima and Haematobia exigua which are serious pests of cattle in Australia.

The Dermestidae are often used in taxidermy and in the preparation of scientific specimens, to clean soft tissue from bones. Beetles are the most widely eaten insects, with about species used as food, usually at the larval stage.

Due to their habitat specificity, many species of beetles have been suggested as suitable as indicators, their presence, numbers, or absence providing a measure of habitat quality.

Predatory beetles such as the tiger beetles Cicindelidae have found scientific use as an indicator taxon for measuring regional patterns of biodiversity.

They are suitable for this as their taxonomy is stable; their life history is well described; they are large and simple to observe when visiting a site; they occur around the world in many habitats, with species specialised to particular habitats; and their occurrence by species accurately indicates other species, both vertebrate and invertebrate.

Many beetles have beautiful and durable elytra that have been used as material in arts, with beetlewing the best example. Whole beetles, either as-is or encased in clear plastic, are made into objects ranging from cheap souvenirs such as key chains to expensive fine-art jewellery.

Monitoring and maintaining proper air pressure in your tires helps with efficiency and safety. The TPMS helps alert you to a loss of tire pressure so you know when to add air.

From Electronic Brake-pressure Distribution EBD , which helps maintain appropriate stopping power during a hard-braking situation, to Electronic Stability Control ESC , which can adjust engine throttle and apply corrective forces to the wheels that need them most, your vehicle is equipped with a total of seven stability-enhancing systems.

Enter your zip code for regional offers. A classic that keeps up with the times. The Beetle says so long with two grand finales. Help yourself to a little extra help.

Beetle SE with Premium Package shown. Color for display only; may not be available on all trims. A warranty built for two.

Compare our warranty to the runners-up. Design The original original. Find one near you. Leather seats with diamond-stitched design.

Leather-wrapped, multi-function steering wheel. Customizable interior ambient lighting. An ever-present reflection of the past.

Uniting classic styling and modern technology, the Beetle has always been ahead of its time. These smartly designed front and rear bumpers add a sporty and modern touch to the Beetle.

Color inside the lines. Performance Feel more in turbocharge. The icon gets a bump in speed. Bitten by the performance bug.

And it gets an EPA-estimated 33 hwy mpg. Technology Up close and personalized. Take your apps for a ride. Learn more about App-Connect Start Over. As entertaining as the curves ahead.

Keyless access with push-button start. Sunroof rain notification Open vehicle notification Parking information. It can send you a notification so you can return to the car and close it.

Driver Assistance features can help give you more confidence on your drive. Features available on select trims. Safety A lot goes in before you do.

But we can engineer. Rear View Camera System. Automatic Post-Collision Braking System. Your mileage will vary and depends on several factors, including your driving habits and vehicle condition.

The Type 32 was similar in design to the Type 12, but it had a flat-four engine. NSU discontinued car manufacturing, and the Type 32 was abandoned at the prototype stage.

In October , the first two Type prototypes, known as cars V1 , a sedan, and V2 , a convertible V for Versuchswagen , or "test car" , were ready.

All cars had the distinctive round shape and the air-cooled , rear-mounted engine. Included in this batch was a rollback soft top called the Cabrio Limousine.

The car was designed to be as simple as possible mechanically. The suspension design used compact torsion bars instead of coil or leaf springs. The Beetle is nearly airtight and will briefly float.

On 26 May , Hitler laid the cornerstone for the Volkswagen factory in Fallersleben. The model village of Stadt des KdF-Wagens was created near Fallersleben in Lower Saxony in for the benefit of the workers at the newly built factory.

Volkswagen had only just started small scale production, building about Beetles, when civilian production was halted at the start of the war. Hitler was given the very first convertible Beetle built in A handful of KdF-Wagens were produced, primarily for the Nazi elite from to , as the Typ It produced small numbers of Kommandeurswagen Typ 87 , with a Typ 1 body mounted on a four-wheel drive Schwimmwagen chassis; the fenders were widened to accommodate Kronprinz all-terrain tires reminiscent of the later Baja Bugs.

Much of the essential equipment had already been moved to underground bunkers for protection, which let production resume quickly after hostilities ended.

Due to gasoline shortages late in the war, a few " Holzbrenner " Beetles were built, powered by pyrolysis gas producers located under the front hood.

In occupied Germany, the Allies followed the Morgenthau plan to remove all German war potential by complete or partial pastoralization.

As part of this, in the Industrial plans for Germany , the rules for which industry Germany was to be allowed to retain were set out.

Mass production of civilian VW cars did not start until post-war occupation. The Volkswagen factory was handed over by the Americans to British control in ; it was to be dismantled and shipped to Britain.

In March , Herbert Hoover helped change policy by stating. There is the illusion that the New Germany left after the annexations can be reduced to a " pastoral state ".

It cannot be done unless we exterminate or move 25,, people out of it. The engine fires up immediately without a choke. It has tolerable road-handling and is economical to maintain.

Although a small car, the engine has great elasticity and gave the feeling of better output than its small nominal size.

There were other, less-numerous models, as well. The Type 18A , a fixed-top cabriolet, was produced by Austro-Tatra as a police and fire unit; were assembled between January and March On 17 February Beetle No.

Though extremely successful in the s, experiencing its greatest sales growth in North America between and , the Beetle was increasingly faced with stiff competition from more modern designs globally.

The Japanese had refined rear-wheel-drive, water-cooled, front-engine, small cars including the Datsun and Toyota Corona , whose sales in the North American market grew rapidly at the expense of Volkswagen in the late s.

Honda introduced the N , based on the space-efficient transverse-engine, front-wheel-drive layout of the original Austin Mini , to the North American market in late , and upgraded the model to the Honda Civic in The Japanese "big three" would soon dominate compact auto sales in North America.

In Ford introduced its Pinto , which had some market impact as a low cost alternative in the wake of the drop of the US Dollar against the Deutschmark that same year.

As the s came to a close, Volkswagen faced increasingly stiff competition from European cars as well. The Beetle was faced with competition from new designs like the Fiat and Renault 5 , and more robust designs based on the Austin Mini layout such as the Superminis.

German competitors, Ford and Opel also enjoyed strong sales of modern smaller cars like the Ford Escort and Opel Kadett. Safety issues with the Beetle came under increasing scrutiny, culminating in the release of a rather scathing report.

None of these models, aimed at more upscale markets, achieved the level of success of the Beetle. The over-reliance on a single model, now in decline, meant that Volkswagen was in financial crisis by Production lines at Wolfsburg switched to the new water-cooled, front-engined, front-wheel-drive Golf designed by Giorgetto Giugiaro in , sold in North America at the time as the "Rabbit".

It was periodically redesigned over its lifetime, with only a few components carried over between generations, entering its seventh generation in ; the Beetle had only minor refinements of its original design.

The Golf did not kill Beetle production, nor did the smaller Polo which was launched a year later. Production of the Beetle continued in smaller numbers at other German factories until 19 January , when mainstream production shifted to Brazil and Mexico: After this shift in production, sales in Europe did not stop, but became very low.

Beetle sedans were produced for U. The last Beetle was produced in Puebla , Mexico, in July Production in Brazil ended in , then started again in and continued until The Beetle outlasted most other cars which had adopted the rear-engine, air-cooled layout such as those by Subaru , Fiat , and General Motors.

By , over 21 million Type 1s had been produced, but by , annual production had dropped to 30, from a peak of 1.

Each car included the 1. A mariachi band serenaded production of the last car. In Mexico, there was an advertising campaign as a goodbye for the Beetle.

In one of the ads was a very small parking space on the street, and many big cars tried to use it, but could not. After a while, a sign appears in that parking space saying: Another depicted the rear end of a Beetle the year Volkswagen was established in Mexico in the left side of the ad, reading " Erase una vez Other ads also had the same nostalgic tone.

In , Volkswagen prototyped a 1. Volkswagen made only two of these naturally aspirated, air-cooled boxer diesel engines, and installed one engine in a Type 1 and another in a Type 2.

The Beetle featured a rear-located, rear-wheel drive, air-cooled four-cylinder, boxer engine in a two-door bodywork featuring a flat front windscreen, accommodating four passengers and providing luggage storage under the front bonnet and behind the rear seat—and offering a coefficient of drag of 0.

The bodywork attached with eighteen bolts to its nearly flat platform chassis which featured a central structural tunnel.

Front and rear suspension featured torsion bars along with front stabilizer bar—providing independent suspension at all wheels.

Certain initial features were subsequently revised, including mechanical drum brakes, split-window rear windows, mechanical direction-indicators and the non-synchronized gearbox.

Other features, including its distinctive overall shape, endured. In fact, the Beetle was prized for its seemingly unchanged appearance and "marketed to American consumers as the anti-GM and Ford: Its engine, transmission, and cylinder heads were constructed of light alloy.

Later models of the carburetor featured an automatic choke. Engine intake air passed through a metallic filter, while heavier particles were captured by an oil bath.

After , steering featured a hydraulic damper that absorbed steering irregularities. Throughout its production, VW marketed the Beetle with a four-speed manual transmission.

From and almost exclusively in Europe , [54] VW offered an optional version of the Saxomat semi-automatic transmission: Subsequently, beginning in in Europe and in the United States , VW offered an optional semi-automatic transmission marketed as Automatic Stick Shift and also called AutoStick , [ citation needed ] which was a 3-speed manual coupled to an electro-pneumatic clutch and torque converter.

It was in that Wilhelm Karmann first bought a VW Beetle sedan and converted it into a four-seated convertible. The convertible was more than a Beetle with a folding top.

To compensate for the strength lost in removing the roof, the sills were reinforced with welded U-channel rails, a transverse beam was fitted below the front edge of the rear seat cushion, and the side cowl-panels below the instrument panel were double-wall.

In addition, the lower corners of the door apertures had welded-in curved gussets, and the doors had secondary alignment wedges at the B-pillar.

The top was cabriolet-style with a full inner headliner hiding the folding mechanism and crossbars. The rear window was tempered safety glass, and after , heated.

Due to the thickness of the top, it remained quite tall when folded. To enable the driver to see over the lowered top, the inside rearview was mounted on an offset pivot.

The convertible was generally more lavishly equipped than the sedan with dual rear ashtrays, twin map pockets, a visor vanity mirror on the passenger side, rear stone shields, and through , wheel trim rings.

Many of these items did not become available on other Beetles until the advent of the optional "L" Luxus Package of After a number of stylistic and technical alterations made to the Karmann cabriolet, [57] corresponding to the many changes VW made to the Beetle throughout its history , the last of , cabriolets came off the production line on 10 January During this period, a myriad of changes were made throughout the vehicle beginning with the availability of hydraulic brakes and a folding fabric sunroof in The rear window of the VW Beetle evolved from a divided or "split" oval, to a singular oval.

The change occurred between October and March The brake light function was transferred to new heart-shaped lamps located in the top of the taillight housings.

In , the separate brake lights were discontinued and were combined into a new larger taillight housing. The traditional VW semaphore turn signals were replaced by conventional flashing directional indicator lamps for North America.

For , the Beetle received what would become one of its more distinctive features, a set of twin chrome tailpipes. Models for North America gained taller bumper guards and tubular overrider bars.

For , the Beetle received a revised instrument panel, and a larger rectangular rear window replaced the previous oval design.

For , significant technical advances occurred in the form of a new engine and transmission. The single-barrel Solex carburetor received an electric automatic choke and the transmission was now synchronized on all forward gears.

The traditional semaphore turn signals were replaced by conventional flashing directional indicators worldwide. For , the Beetle received a mechanical fuel level gauge in place of the former fuel tap.

The Standard model continued without a gas gauge until the end of the model year. At the rear, larger tail lights were introduced incorporating a separate amber turn signal section to meet new European standards these turn signals remained red in the US market until The former hand-pump style windscreen washer was replaced by a new design using compressed air.

The largest change to date for the Beetle was in The result was a more open, airy, modern look. Models so equipped carried a "" badge on the engine lid.

The cc engine was standard for North America. For , a yet-again larger- displacement engine was made available: North America received the engine as standard equipment, but did not receive front disc brakes.

These models were identified by a "Volkswagen" badge on the engine lid. The rear suspension was significantly revised and included a widened track, softer torsion bars and the addition of a unique Z-configuration equalizing torsion spring.

The clutch disc also increased in size and changes were made to the flywheel. In accord with the newly enacted US Federal Motor Vehicle Safety Standard , North American models received a dual-circuit brake system, the clear glass headlamp covers were deleted; the headlamps were brought forward to the leading edge of the front fenders, and the sealed-beam units were exposed and surrounded by chrome bezels.

In the rest of the world markets the model retained the older headlights. Another oddity of the North American market Beetle is the rear bumper overriders towel rails — the overriders have a different shape than the older models besides the one-year only engine decklid.

The most noticeable of which were the new larger, higher mounted C-section bumpers. At the rear, new larger taillamps were adopted and were able to accommodate reversing lamps, which were previously separate bumper-mounted units.